Undergraduate Course Catalog

UWM Undergraduate Catalog 2017-2018

College of Letters and Science

Biological Sciences

Biology is the study of life. Biologists analyze organisms at the cellular and molecular levels using genetics, biochemistry, and microscopy. They also study interactions of organisms with each other and with the environment using behavior, morphology, and genetics. These studies have applications across many areas including agriculture, medicine, and the environment.

Majors in biology are needed in areas such as farming, food processing, bioremediation, as well as the biomedical and biotechnology fields. A biology major also prepares students for professional careers in medicine, dentistry, pharmacy, veterinary science, and education, in addition to advanced studies in any life science field.

The Department offers two majors: Biological Sciences and Microbiology. The Biological Sciences major has two options: the Standard option and the Cell and Molecular Biology option. The Microbiology major has two options: the Standard option and the Industrial Fermentation and Biotechnology option. Both majors and options are excellent preparation for a career in a life science field. The multiple offerings allow students a broad choice in their studies.

Course of Study: Biological Sciences Major

The Biological Sciences major requires a minimum of 34 credits in Biology, of which 26 must be at the advanced (300 and above) level. At least 15 of the advanced credits must be taken in residence at UWM. Students must complete four (4) laboratory courses. No more than eight (8) credits in 100-level courses in Biological Sciences may be applied toward the major, and students may not combine Bio Sci 150, 202, and 203 for more than 9 credits toward the major.�A combined limit of 6 credits in Bio Sci 290, 489, 695, 697, 698, 699, Honors 686, 687, and 689 counts toward the major.�Students must attain a GPA of 2.500 in Biological Sciences courses attempted at UWM. In addition, students must attain a GPA of at least 2.500 in all major credits attempted, including any transfer work.

Additional requirements for the major include the following foundational courses in Math, Physics, and Chemistry: 1) Math 211, 221, or 231 for a Bachelor of Science (Math 105 or equivalent for a Bachelor of Arts); 2) One year of Physics (120/122 or 209/210, with at least one semester of Physics lab (121 or 214) or one year of Physics in the studio format (219/220); and 3) Either the survey of Organic Chemistry with lab (Chem 341/342) or the one-year Organic Chemistry sequence with one lab (Chem 343/344/345). For students in the Cell and Molecular Biology (CMB) option, Chem 343, 344, and 345 are required, and a second physics lab is strongly recommended. Students who plan to attend graduate or professional schools are advised to take the one-year sequence of Organic Chemistry with lab, a course in Biochemistry (Chem 501; also counts toward major), and two semesters of Calculus.

STANDARD BIOLOGICAL SCIENCES OPTION

Required Biological Sciences Courses
Bio Sci 150	Foundations of Biological Sciences I	4
Bio Sci 152	Foundations of Biological Sciences II	4
Bio Sci 325	Genetics	4
Either:
Bio Sci 310	General Ecology	4
or both
Bio Sci 315 and	Cell Biology	3
Bio Sci 316	Laboratory in Genetics and Cell Biology	2
Research Requirement: One of Bio Sci 611, 670, 671, 672, or CES 490 (a senior seminar); Honors 686, 687, or 689 (Honors research, project, or thesis); or Bio Sci 695, 697, 698, or 699 (Independent Study).
Electives to reach a total of 26 upper-division (numbered 300 and above) credits and one additional laboratory course.

CELL AND MOLECULAR BIOLOGY OPTION

Core Courses
Bio Sci 150	Foundations of Biological Sciences I	4
Bio Sci 152	Foundations of Biological Sciences II	4
Bio Sci 315	Cell Biology	3
Bio Sci 316	Laboratory in Genetics and Cell Biology	2
Bio Sci 325	Genetics	4
Laboratory Courses: At least one Cell and Molecular Biology laboratory numbered above 350 or BIO SCI 203. See individual course descriptions to identify eligible courses.
Electives: Any U-only Cell and Molecular Biology course numbered between 350 and 399.
Any U/G Cell and Molecular Biology course numbered 400 or above.
Research Requirement: Either BIO SCI 672 (Undergraduate Seminar in Cell and Molecular Biology) or BIO SCI 697 (Independent Study in Cell and Molecular Biology).

Students must select additional Cell and Molecular Biology courses to reach a total of 18 credits beyond the core courses and a total of 26 upper-division (numbered 300 and above) credits.

Course of Study: Microbiology Major

Microbiology is the study of microorganisms such as bacteria, archaea, protists, fungi, and viruses. Prospective Microbiology majors should consult with a faculty advisor as early as possible, preferably before the beginning of the junior year, in order to outline an appropriate course of study. Students should consult their advisor at least once each semester.

There are two options within the Microbiology major: the Microbiology Standard option and the Industrial Fermentation and Biotechnology option, which is offered jointly with the Department of Chemistry and Biochemistry. For both options, at least 15 credits of advanced (300 and above) Microbiology courses must be taken in residence at UWM. Students must attain an average GPA of 2.500 in Microbiology courses attempted at UWM. In addition, students must attain a 2.500 GPA on all major credits attempted, including any transfer work.

Microbiology Standard Option
The Microbiology option requires a minimum of 34 credits in Microbiology. The required and elective courses in Bio Sci, Chem 501, and BMS 534, 535, 539, and 540 count as "Microbiology" courses for this purpose. A combined limit of 6 credits in Bio Sci 290, 695, 697, 698, and 699 counts toward the major.

Bio Sci 150	Foundations of Biological Sciences I	4
Bio Sci 152	Foundations of Biological Sciences II	4
Bio Sci 325	Genetics	4
Bio Sci 383	General Microbiology	4
Bio Sci 529	Molecular Biology of Microorganisms	3
Bio Sci 540	Microbial Diversity and Physiology	3
Research Requirement: Bio Sci 495, 671, 698 or, when determined by the student's Microbiology faculty advisor to have Microbiology content, Bio Sci 699, or Honors 686, 687, or 689. (For titles and credits, see electives list.)
Elective Microbiology Courses: Students choose courses from this list to complete their 26 credits of upper-level (300 or above) microbiology courses. Students MUST take EITHER Bio Sci 315 (Cell Biology) OR Chem 501 (Introduction to Biochemistry), and they MUST take EITHER Bio Sci 537 (Industrial Microbiology and Biochemistry Laboratory), OR Bio Sci 539 (Laboratory Techniques in Molecular Biology), OR Bio Sci 580 (Experimental Microbiology).
Bio Sci 315	Cell Biology	3
Bio Sci 316	Laboratory in Genetics and Cell Biology	2
Bio Sci 401	Immunology	2
Bio Sci 402	Immunological Techniques	3
Bio Sci 405	General Virology	3
Bio Sci 425	Plankton Biology	4
Bio Sci 490	Molecular Genetics	3
Bio Sci 495	Internship in Biotechnology, Upper Division	3-6
Bio Sci 507	Environmental Microbiology	3
Bio Sci 535	Bacterial Pathogenesis	3
Bio Sci 536	Applied Microbiology and Biotechnology	2
Bio Sci 537	Industrial Microbiology and Biochemistry Laboratory	2
Bio Sci 539	Laboratory Techniques in Molecular Biology (if not selected above)	4
Bio Sci 542	Biological Electron Microscopy	3
Bio Sci 544	Transmission Electron Microscopy Laboratory	3
Bio Sci 572	Genomics	3
Bio Sci 573	Cellular Evolution	3
Bio Sci 580	Experimental Microbiology	4
Bio Sci 595	Principles of Genetic and Molecular Engineering	3
Bio Sci 597	RNA Structure, Function, and Metabolism	3
Bio Sci 667	Advanced Techniques in Microbial, Molecular, and Cellular Biology	4
Bio Sci 671	Undergraduate Seminar in Microbiology: (subtitle) (may not count for more than 1 of the elective credits)	1
Bio Sci 698	Independent Study in Microbiology	1-3 (per semester)
Bio Sci 699¹	Independent Study	1-3 (per semester)
Chem 501	Introduction to Biochemistry	3
Chem 601	Biochemistry: Protein Structure and Function	3
BMS 534²	Medical Microbiology	2
BMS 535²	Medical Microbiology Laboratory	2
BMS 539²	Public Health Microbiology	2
BMS 540²	Public Health Microbiology Laboratory	1
Honors 686¹	Research in Honors	2-3
Honors 687¹	Senior Honors Project	1-6
Honors 689¹	Senior Honors Thesis	3

¹ Only Bio Sci 699 or Honors 686, 687, or 689 projects that are determined by the student's microbiology faculty advisor to have microbiology content count toward the major.

² Students may elect to take these BMS courses to earn credit toward the Microbiology major option provided that they earn 30 advanced credits in Natural Sciences within the College of Letters and Science, as required for the BS degree.

Other Required Courses
Chem 102	General Chemistry	5
Chem 104	General Chemistry and Qualitative Analysis	5
Chem 343	Organic Chemistry	3
Chem 344	Organic Chemistry Laboratory	2
Chem 345	Organic Chemistry	3
Math 211 or	Survey in Calculus and Analytic Geometry	4
Math 221 or	Honors Calculus I	5
Math 231	Calculus and Analytical Geometry I	4
AND
Math 222 or	Honors Calculus II	5
Math 232 or	Calculus and Analytic Geometry II	4
Bio Sci 465	Biostatistics	3
One of the following sets of physics courses:
Physics 120	General Physics I (Non-calculus Treatment)	4
Physics 122	General Physics II (Non-calculus Treatment)	4
Physics 123	General Physics Laboratory II (Non-calculus Treatment)	1
or

Physics 209	Physics I (Calculus Treatment)	4
Physics 210	Physics II (Calculus Treatment)	4
Physics 215	Lab Physics II (Calculus Treatment)	1
or

Physics 219	Physics I: Calculus-Based, Studio Format	5
Physics 220	Physics II: Calculus-Based, Studio Format	5

Industrial Fermentation and Biotechnology Option

Bio Sci 150	Foundations of Biological Sciences I	4
Bio Sci 152	Foundations of Biological Sciences II	4
Bio Sci 325	Genetics	4
Bio Sci 383	General Microbiology	4
Bio Sci 529	Molecular Biology of Microorganisms	3
Bio Sci 536	Applied Microbiology and Biotechnology	3
Bio Sci 537	Industrial Microbiology and Biochemistry Laboratory	2
Chem 501	Introduction to Biochemistry	3
Chem 541	Bioprocess Chemical Engineering	3
Chem 543	Bioproduct Regulatory Protocols	3
Chem 690	Seminar: Science Career Transitions	1
One of
Bio Sci 489 or	Internship in Biological Sciences, Upper Division (Industry Internship)
Bio Sci 698	Independent Study in Microbiology
Other Required Courses
Chem 102	General Chemistry	5
Chem 104	General Chemistry and Qualitative Analysis	5
Chem 343	Organic Chemistry	3
Chem 344	Organic Chemistry Laboratory	2
Chem 345	Organic Chemistry	3
One of
Math 211 or	Survey in Calculus and Analytic Geometry	4
Math 221 or	Honors Calculus I	5
Math 231	Calculus and Analytical Geometry	4
One of
Math 222 or	Honors Calculus II	5
Math 232 or	Calculus and Analytical Geometry	4
Bio Sci 465	Biostatistics	3
One of the following sets of Physics courses:
Physics 120	General Physics I (Non-calculus Treatment)	4
Physics 122	General Physics II (Non-calculus Treatment)	4
Physics 123	General Physics Laboratory II (Non-calculus Treatment)	1
or
Physics 209	Physics I (Calculus Treatment)	4
Physics 210	Physics II (Calculus Treatment)	4
Physics 215	Lab Physics (Calculus Treatment)	1
or
Physics 219	Physics I: Calculus-Based, Studio Format	5
Physics 220	Physics II: Calculus-Based, Studio Format	5

Honors in the Major

Students in biological sciences who meet all of the following criteria are awarded honors in the major upon graduation:

3.500 cumulative GPA in all UWM graded credits attempted;
3.750 GPA in UWM courses counting toward the major;
3.500 GPA in all advanced credits that count toward the major; and
Complete a laboratory or field research independent study (Bio Sci 697, 698, or 699) or internship (Bio Sci 489 or CES 489).

Students who believe they may qualify for honors in biological sciences should apply to the Department during their last semester of study.

Course of Study: Minor

Students with a major in another discipline can apply for a minor in Biological Sciences by meeting the requirements listed below, completing the appropriate “Declaration of Minor” form, and meeting with a Department of Biological Sciences advisor.

Requirements: The minor consists of 19 credits in Biological Sciences including the core introductory sequence (Bio Sci 150 and Bio Sci 152); a gateway course selected from the following three options 1) Bio Sci 310, or 2) Bio Sci 315 plus Bio Sci 316, or 3) Bio Sci 325 plus Bio Sci 316; and at least 7 additional credits in courses that carry credit toward the major. At least 9 credits must be taken at or above the 300 level in residence at UWM. Students must maintain a GPA of 2.500 in minor courses attempted at UWM. In addition, students must attain a GPA of 2.500 in all minor courses attempted, including any transfer work.

Declaration of Major or Minor in Biological Sciences
Students should contact an advisor (see list below) as soon as possible in their freshman year about required courses and the recommended course sequence within Biological Sciences. They should complete Bio Sci 150 and be enrolled in Bio Sci 152 before officially declaring a major in Biological Sciences. Upon declaring the major, the student must select a Department of Biological Sciences faculty advisor (from the list below) who will guide the student in planning his/her curriculum and will help the student accomplish his/her goals. Failure to complete the declaration of major, as recommended, may result in a delay in graduation.

Procedure for Processing a Declaration of Major or Minor:

1. Obtain and complete a “Declaration of Major (or Minor) Form” from the Department of Biological Sciences office (Lapham Hall S181).

2. Attend the Open Advising session held before registration each semester or select a departmental advisor in the area of interest (see list below) and set up an advising appointment by sending an email to the prospective advisor as follows:

* The “subject box” of the email should say “Appointment to Declare Major (or Minor).”

* The text of the message should give days and times the student is available to meet for advice about the major/minor and curriculum. Take the “Declaration of Major/Minor Form” and an up-to-date copy of the student's transcript (available from PAWS) to the advising appointment at which the advisor will sign the form. After this initial meeting, return to meet with the departmental advisor regularly to discuss academic progress and/or attend open advising regularly.

3. Return the "Declaration of Major/Minor Form" with the advisor's signature, to Lapham Hall S181 to obtain the Department Chair's signature. The Chair will forward the form to the College of Letters and Science Office of Student Academic Services.

4. Meet each semester with the assigned College of Letters and Science advisor (in Holton Hall); the L&S advisor will monitor progress toward completion of the L&S degree requirements.

Department Advisors

Biological Sciences Advisors	Lapham Office	E-mail (@uwm.edu)
Professor Berges – Marine Biology	S587	berges
Professor Coggins – Parasitology	S297	coggins
Professor Dunn – Behavioral Ecology	S497	pdunn
Professor Karron – Plant Ecology	S585	karron
Professor Latch – Population Genetics	N215	latch
Professor Rodr�guez – Behavioral Ecology	S295	rafa
Professor Strickler – Ecology	Water Inst. 137	jrs
Professor Whittingham – Behavioral Ecology	S499	whitting
Professor Young – Plant Biology	S593	ebyoung
Cellular & Molecular Biology Option Advisors
Professor Heathcote – Neuroscience	N411	rdh
Professor Hutz – Physiology	N509	rjhutz
Professor Oliver – Mammalian Cell Biology	N209	oliver
Professor Scanes – Animal Physiology & Nutrition	S493	scanes
Professor Steeber – Immunology	N211	steeber
Professor Udvadia – Neuroscience	SB80	audvadia
Professor Wejksnora – Molecular Biology	220	pjw
Professor Wimpee – Molecular Biology	S495	cwimpee
Professor Witten – Neuroscience	N409	jlw
Professor Zhao – Molecular Genetics	462	dzhao
Microbiology Advisors
Professor Dey – Molecular Biology	460	deym
Professor Forst – Molecular Biology	458	sforst
Professor Kuchin – Genetics	444	skuchin
Professor McBride – Molecular Biology	N307	mcbride
Professor Prasad – Microbiology	440	prasadg
Professor Saffarini – Environmental Microbiology	N309	daads
Professor Yang – Genomics	131D	chyang
Advisors for Related Programs
Major in Conservation & Environmental Science
Professor Fredlund – Biogeography	364	fredlund
Pre-Professional (Medicine, Dental, Pharmacy, Veterinary Medicine)
Patty Cobb (pre-Medicine)	Holton 131	pacobb
Jenny Hack (pre-Dentistry)	Holton 150	jlrahmer
Laura Stark (pre-Pharmacy)	Holton 130	starll
Lori Fitzenberger (pre-Veterinary Medicine)	Holton 147	lorifitz

Related Areas of Study

The Conservation and Environmental Science major is an alternative, interdisciplinary program for students with specific interests in conservation or environmental science (see Interdepartmental Majors, College of Letters and Science).

For the Pre-Forestry (Wildlife Management) curriculum, see the Pre-Professional Programs section (College of Letters and Science).

Pigeon Lake Field Station is a natural laboratory sponsored by the 14 campuses of the University of Wisconsin System, located in the Chequamegon National Forest, 30 miles from Lake Superior in northern Wisconsin's Bayfield County. Pigeon Lake Field Station offers summer programs in a variety of disciplines, including biology. Work completed at Pigeon Lake is credited as resident study by UWM. For additional information, contact the Biological Sciences Department office.

The UWM Field Station is an area of almost 2,500 acres devoted to the study of natural history and biology. Located about 25 miles north of campus on the Cedarburg Bog in the Town of Saukville, the station includes a wide variety of natural areas ranging from old-growth forest to acid bog and several lakes. Many biological sciences courses use the Field Station for instruction and to provide hands-on research opportunities. The Field Station provides internship and independent study opportunities for biology students; it also offers short courses for credit through the department. For additional information, contact the Biological Sciences Department office.

Courses

Biological Sciences (BIO SCI)

Cross-listed Courses

The following courses offered by other departments may be used to fulfill the requirements of the undergraduate major and minor in biological sciences:

CES 471	Practicum in Natural Resource Management
CES 490	Senior Seminar: Conservation and Environmental Sciences
Chem 501	Introduction to Biochemistry
Psych 254	Physiological Psychology
Psych 654	Advanced Physiological Psychology

Faculty and Staff

Filipe Alberto, Asst. Prof., PhD
University of Algarve, Portugal

Sonia Bardy, Asst. Prof., PhD
Queen's University, Ontario

John Berges, Prof., PhD
University of British Columbia

Gerald Bergtrom, Prof. Emeritus, PhD

Martin Boraas, Assoc. Prof. Emeritus, PhD

Arthur S. Brooks, Prof. Emeritus, PhD

John Buntin, Prof. Emeritus, PhD

James R. Coggins, Assoc. Prof. Emeritus, PhD

Mary Lynne Perille Collins, Prof. Emerita, PhD

Madhusudan Dey, Assoc. Prof., PhD
Jawaharlal Nehru University, India

Peter Dunn, Prof., PhD
University of Alberta

Timothy Ehlinger, Assoc. Prof., PhD
Michigan State University

Millicent S. Ficken, Prof. Emerita, PhD

Steven Forst, Prof., PhD
New York University

Ralph Grunewald, Assoc. Prof. Emeritus, PhD

Jennifer Gutzman, Asst. Prof., PhD
University of Wisconsin-Madison

R. David Heathcote, Prof., PhD, Chair
University of California, Berkeley

Gerlinde Hoebel, Asst. Prof., DNS
University of Ulm, Germany

Sara B. Hoot, Prof. Emerita, PhD

Reinhold Hutz, Prof., PhD
Michigan State University

Jeffrey Karron, Assoc. Prof., PhD
University of Colorado

Geoffrey S. Kennedy, Assoc. Prof. Emeritus

Sergei Kuchin, Assoc. Prof., PhD
Institute of Genetics (Moscow)

Emily Latch, Assoc. Prof., PhD
Purdue University

Jakob R. Loewenberg, Prof. Emeritus, PhD

Mark McBride, Prof., PhD
University of Wisconsin-Madison

Julie A. Oliver, Assoc. Prof., PhD
University of Wisconsin-Madison

Marianna Orlova, Instrument Innovator, PhD
Institute of Genetics and Selection of Industrial Microorganisms, Moscow

Heather A. Owen, Assoc. Scientist, PhD
Miami University

Andrew Petto, Lect., PhD
University of Massachusetts, Amherst

Ruth B. Phillips, Prof. Emerita, PhD

Gyaneshwar Prasad, Assoc. Prof., PhD
Maharaja Sayajirao University of Baroda, India

Christopher C. Quinn, Asst. Prof., PhD
Yale University

Charles C. Remsen, Prof. Emeritus, PhD

Rafael Rodríguez, Assoc. Prof., PhD
University of Kansas

Daad A. Saffarini, Prof., PhD
University of Wisconsin-Milwaukee

Colin Scanes, Prof. Emeritus, PhD, DSc

Dianne Seale, Assoc. Prof. Emerita, PhD

Cynthia V. Sommer, Assoc. Prof. Emerita, PhD

Douglas Steeber, Assoc. Prof., PhD
University of Wisconsin-Madison

J. Rudi Strickler, Distinguished Prof., PhD
Swiss Federal Institute of Technology

Ava Udvadia, Assoc. Prof., PhD
Duke University

Donna Van Wynsberghe, Prof. Emerita, PhD

Eldon D. Warner, Prof. Emeritus, PhD

Roslyn P. Warren, Prof. Emerita, PhD

Peter J. Wejksnora, Assoc. Prof., PhD
Brandeis University

Linda A. Whittingham, Prof., PhD
Queens University, Ontario

Charles F. Wimpee, Assoc. Prof., PhD
University of California, Los Angeles

Jane Witten, Assoc. Prof., PhD
University of Chicago

Ching-Hong Yang, Prof., PhD
University of California, Riverside

Erica Young, Assoc. Prof., PhD
Monash University (Melbourne, Australia)

Dazhong Zhao, Assoc. Prof., PhD
Chinese Academy of Science

Adjunct Faculty

Gretchen A. Meyer, Adjunct Senior Scientist, PhD
Cornell University

James R. Moyer, Jr., Assoc. Prof., PhD, Psychology
Northwestern University

Valerica Raicu, Prof., PhD, Physics
University of Bucharest, Romania

James A. Reinartz, Adjunct Senior Scientist, PhD
Duke University

Marius Schmidt, Prof., PhD, Physics
Technical University of Munich

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University of Wisconsin-Milwaukee Undergraduate Catalog 2017-2018:
Biological Sciences
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